Jack Billabard, mourning the loss of his wife and baby in childbirth, vows to never to love again. After their funeral at Fort Laramie, he rides into the Wyoming hills beyond the ranch he built for his wife. Through his grieving tears, an ancient tree appears, giving him the hope he doesn't believe is possible. For the next four years, he acts as a guide on the Oregon Trail, taking families to a new life while his looms lonely and stagnant.

The night before her abusive husband’s death, an ancient tree appears in Sarah Nickelson's yard as she agonizes over how to survive her marriage. The tree gives her hope she can’t help but reject. After all, a tree doesn't just appear out of nowhere. After her husband 's death, and with no options as a widow in Independence, Missouri, Sarah decides to travel to Oregon City as a Mail Order Bride.

During their trek west on the Oregon Trail, Jack and Sarah encounter one another, each afraid of being hurt again. Can they survive dogs and puppies, wind and rainstorms, Indians and unfavorable fellow passengers, while their love blossoms? Will the tree fulfill its promise?